Biology and Biological Engineering Division
B.S., Caltech, 1955; Ph.D., University of Cambridge, 1958. Visiting Assistant Professor, Caltech, 1960; Assistant Professor, 1961-63; Associate Professor, 1963-68; Professor, 1968-2000; Professor Emeritus, 2000-. Executive Officer for Biology, 1976-80; 1985-89; Associate Chairman of the Division of Biology, 1980-85.