Biology and Biological Engineering Division
B.S., Pennsylvania State University, 1983; Ph.D., Caltech, 1987. Senior Research Fellow, Caltech, 1991-92; Assistant Professor, 1992-98; Associate Professor, 1998-2003; Professor, 2003-07; Bren Professor, 2007-; Merkin Institute Professor, 2021-; HHMI Investigator, 1994-07; Executive Officer for Biochemistry and Molecular Biophysics, 2004-07; Vice Provost, 2007-10; Division Chair, 2010-20; Bowes Jr. Leadership Chair, 2012-20.